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The antiparallel orientation of complementary strands in duplex DNA was elegantly demonstrated in 1960 by Arthur Kornberg by nearest-neighbor analysis. In this technique, DNA is synthesized by DNA polymerase I from one deoxynucleoside triphosphate whose α-phosphate group is radioactively labeled with ^32 P and three unlabeled deoxynucleoside triphosphates. The resulting product is treated with an enzyme, DNase I, that catalyzes the hydrolysis of the phosphodiester bonds on the 3^' sides of all d
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